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However, based on skull morphology in 1994, it was proposed that I. g. boliviensis was a different species. In 2002, following the analysis of mitochondrial DNA specimens from the Orinoco basin, the Putumayo River (tributary of the Amazon) and the Tijamuchy and Ipurupuru rivers, geneticists proposed that genus Inia be divided into at least two evolutionary lineages: one restricted to the river...